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Professor of Internal Medicine, University of Virginia

PhD in Biochemistry from the University of California, Berkeley

Postdoctoral research at MIT.

I direct a laboratory at the University of Virginia that is funded by the NIH (NHLBI and NIA), NASA and an upcoming grant from the Department of Defense.

Our work is focused on genome instability (somatic mosaicism) and how this contributes to aging and age-associated diseases.

We have published over 400 articles (h index: 131 Google scholar; 111 Scopus).


  • –present
    Professor, University of Virginia


  • 1984 
    University of California, Berkeley, PhD