Toby Tyrrell is professor of Earth system science at the National Oceanography Centre Southampton, University of Southampton. He specialises in biogeochemical cycles.
How organisms interact with their environments
Ocean biogeochemistry, both modern and through Earth history
Ocean chemical changes during extreme events in Earth’s ancient past (for instance at the E/O and K/T boundaries)
Marine cycles of N, P, C, Si
Ecology of phytoplankton, particularly coccolithophores
Controls on biogenic element distributions in the sea, whether from ecological competition between phytoplankton functional groups or from other causes
How Earth stayed habitable for 3 billion years