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A double major in Zoology and Physiology at UWA in the 1960s determined my future path in trying to fuse these two disciplines in an ecological context and develop what is now known as ecophysiology.

Various postdoctoral positions in the UK, France and the United States added skills in endocrinology, which were needed to respond to the question of how animals cope physiologically with, and resolve, the many competing stresses in their natural environment.

My teaching and research has been spread across all the vertebrate animal groups, with adaptation (or in some cases the lack of it) as a primary theme. Since retiring from the Foundation Chair of Zoology in 2005, I have focused on the major concern confronting us today – the protection and conservation of what remains of our natural environment.


  • –present
    Emeritus Professor and Senior Honorary Research Fellow, University of WA


Kelvin Gold Medal of the Royal Society of Western Australia 2010; Awarded a Special Commendation Whitley Award for 2015 of the Royal Zoological Society of New South Wales (RZSNSW) for the promotion of knowledge and conservation of Australasian fauna through many outstanding publications over an extended time period.