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Professor of Forest Ecology and Physiology, Penn State

B.S., S.U.N.Y. Binghamton (1976)
M.S., Michigan State University (1979)
Ph.D., Michigan State University (1982)
Academic Interests:
Disturbance ecology; old-growth forests; tree physiology; vegetation classification
Affiliated Programs:
Graduate faculty, Intercollege Graduate Degree Program in Ecology and Plant Physiology.
Courses Taught:
Forest Ecology; Tree Physiology
Professional Affiliation:
Editor, Ecology ; Panel Member, National Science Foundation, Division of Environ-mental Biology, Collaborative Research for Undergraduate Institutions (CRUI) Grants; Member, Ecological Society of America Committee on Vegetation Classification


  • 1982–present
    Professor, Penn State University