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Scaling up from the ecology of individual animals and plants to populations, communities and ultimately whole ecosystems is my primary research goal. I try to do this in three ways:

by studying the behaviour of animals and the fate of plants at the scale of individual organisms, and using this information to predict how groups of individuals might perform
by documenting ecosystem-scale patterns in biodiversity, and to link these patterns with important environmental variables like water flow or food supply, and
by making ecological models that try to unify processes acting at different scales into a holistic understanding of ecosystems
Most of this work is set in between the tidemarks on rocky shores and sandy beaches among the habitats and species I know best.

At the Scottish Marine Institute I am a member of the ...

Ecology Department
Dynamic Oceans Theme
Modelling group


  • –present
    Head of Ecology, Scottish Association for Marine Science