BSc, National Taiwan University (2003-2006); MSc, Eidgenossische Technische Hochschule (ETH) Zurich (2007-2008); Dr. rer. nat., Freie Universitat Berlin (2009-2012, P. H. Seeberger). Research Assistant, Academia Sinica (2006-2007, S.-H. Wu); MRC Career Development Fellow and EMBO Fellow, MRC Laboratory of Molecular Biology (2012-2014, J. W. Chin). Appointed Lecturer, Cardiff (2015).